Achillea millefolium subsp. compacta (Lam.) Bonnier & Layens accepted as Aspilia foliacea (Spreng.) Baker (original description)
Anthemis maritima subsp. secundiramea (Biv.) Bonnier & Layens accepted as Anthemis secundiramea Biv. (original description)
Carduus nigrescens subsp. vivariensis (Jord.) Bonnier & Layens (original description)
Centaurea jacea subsp. nigra (L.) Bonnier & Layens accepted as Centaurea nigra L. (original description)
Hieracium lanatum subsp. liottardii (Vill.) Bonnier & Layens accepted as Hieracium tomentosum subsp. liottardii (Vill.) Zahn (original description)
Leucanthemum vulgare subsp. pallens (Perreym.) Bonnier & Layens accepted as Leucanthemum pallens (Perreym.) DC. (original description)
Santolina chamaecyparissus subsp. pectinata (Lag.) Bonnier & Layens accepted as Santolina pectinata Lag. (original description)